Prokaryote, relatively simple unicellular
organism, such as a bacterium, characterized by the absence of a nucleus and
other specialized cell structures. Scientists distinguish prokaryotes from
eukaryotes, which are more complex organisms with cells that contain a nucleus,
such as plants and animals.
Structure
of prokaryotes
Prokaryotic cells are relatively small, ranging in
size from 0.0001 to 0.003 mm (0.000004 to 0.0001 in) in diameter. With the
exception of a few species, prokaryotic cells are surrounded by a protective
cell wall. Just inside the cell wall of prokaryotes is the
plasma membrane, a thin structure that is both flexible and strong. In both
prokaryotes and eukaryotes, the plasma membrane is composed of two layers of
phospholipid molecules interspersed with proteins, and regulates the traffic
that flows in and out of the cell. The prokaryotic plasma membrane, however,
carries out additional functions. It participates in replication of
deoxyribonucleic acid (DNA) for cell division and synthesis of adenosine
triphosphate (ATP), an energy molecule. In some prokaryotes, the plasma
membrane is essential for photosynthesis, the process that uses light energy to
convert carbon dioxide and water to glucose.
Reproduction
of Prokaryotes
Most prokaryotes multiply
by the asexual process of binary fission, in which the DNA of the organism
replicates in the cytoplasm, the cell divides in two, and one DNA molecule
passes to each newly formed cell. In addition, some prokaryotes undergo various
processes of genetic recombination. For example, in the process called
transformation, a bacterium removes one or more genes from one organism and
incorporates the genes into its own genetic makeup. In conjugation two
organisms exchange genes. In transduction a virus transports bacterial genes
from one organism to another. Gene transfers account for the appearance of new
biochemical traits in prokaryotes.
Nutrition
Like most organisms, prokaryotes
require carbon and energy to create nutrients such as carbohydrates, proteins,
lipids, and nucleic acids. Prokaryotes obtain carbon and energy from a variety
of sources. Certain prokaryotes use carbon dioxide as their carbon source.
Called autotrophs, these prokaryotes derive energy from different sources, such
as photosynthesis or inorganic molecules. Photoautotrophs, including the
cyanobacteria and the green sulfur and purple sulfur archaebacteria, derive
their energy from light. Chemoautotrophs, such as the soil bacteria Nitrobacter
and Nitrosomonas, derive their energy from inorganic compounds such as
hydrogen sulfide, ammonia, and iron. Heterotrophs are organisms that rely on
ready-made organic compounds such as glucose or alcohol for their carbon
source. Heterotrophs obtain energy by degrading organic molecules, such as
plant or animal matter. A small group of bacteria, the photoheterotrophs, use
light as their energy source, while chemo heterotrophs use organic compounds
for both their carbon and energy sources.
Importance of Prokaryotes
Prokaryotes play significant
roles in our daily lives. In a process called nitrogen fixation, many species
of cyanobacteria convert atmospheric nitrogen to nitrogenous compounds that
other organisms use as food sources. Moreover, the photosynthesis occurring in
cyanobacteria still contributes substantial amounts of oxygen to the atmosphere
and stores the Sun’s energy in carbohydrate molecules. Cyanobacteria are the
foundation for aquatic ecosystems, providing food for protozoa and other
aquatic organisms. Cyanobacteria are threatened, however, by ultraviolet
radiation, which penetrates the atmosphere as a result of the thinning ozone
layer.
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